1. MOE Key Laboratory of Bioinformatics, School of Life Sciences, Tsinghua University, Beijing 100084, China
2. MOE Key Laboratory of Bioinformatics; Bioinformatics Division and Center for Synthetic & Systems Biology, TNLIST; Department of Automation, Tsinghua University, Beijing 100084, China
3. Bioinformatics Center in School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu 610054, China
4. Center of Bioinformatics (COBI), Key Laboratory for Neuro Information of Ministry of Education, University of Electronic Science and Technology of China, Chengdu 610054, China
Corresponding author: Zhirong Sun
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