¹Research Centre for Deep Sea and Polar Fisheries, and Key Laboratory of Mariculture, Ministry of Education, Ocean University of China, 266003, Qingdao, China

²Frontiers Science Center for Deep Ocean Multispheres and Earth System, Ocean University of China, 266100, Qingdao, China

³Alfred Wegener Institute, Helmholtz Centre for Polar and Marine Research, Am Handelshafen 12, 27570, Bremerhaven, Germany

⁴Helmholtz Institute for Functional Marine Biodiversity at the University Oldenburg, Ammerländer Heerstraße 231, 23129, Oldenburg, Germany

⁵Institute for Marine and Antarctic Studies, University of Tasmania, 7004, Hobart, Australia

⁶Fenner School of Environment and Society, Australian National University, 2602, Canberra, ACT, Australia

⁷College of Fisheries, Ocean University of China, 266003, Qingdao, China

⁸Polar Research Institute of China, 200136, Shanghai, China