¹State Key Laboratory of Microbial Metabolism, School of Oceanography, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, 200240, Shanghai, China

²Shenzhen Key Laboratory of Marine Microbiome Engineering, Institute for Advanced Study, Shenzhen University, 518060, Shenzhen, China

³Key Laboratory of Microbial Resources, Institute of Microbiology Chinese Academy of Sciences, 100101, Beijing, China

⁴College of Marine Life Sciences and Institute of Evolution and Marine Biodiversity, Ocean University of China, 266003, Qingdao, China