A new Late Permian species of Curvicubitidae illuminates the phylogeny and palaobiogeography of Pereborioidea (Insecta: Hemiptera)

Ziqiang Xu; ChungKun Shih; Chenxin Zhou; Wei Yuan; Yu Liu; Dong Ren; Ying Wang

doi:10.1111/jse.70016

Journal of Systematics and Evolution ›› 2026, Vol. 64 ›› Issue (2) :330 -343. DOI: 10.1111/jse.70016

Research Article

A new Late Permian species of Curvicubitidae illuminates the phylogeny and palaobiogeography of Pereborioidea (Insecta: Hemiptera)

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Abstract

A new species, Sinocurvicubitus haotianus Xu, Shih, Ren & Wang, sp. nov. (Curvicubitidae), is described from the Upper Permian Leping Formation in Jiangxi Province, South China. This constitutes the second definitive occurrence of Curvicubitidae in the Late Permian. Through comparative wing morphology and cladistic analyses, we resolved phylogenetic relationships within the superfamily as ((Ignotalidae + Pereboriidae*) + Curvicubitidae) and validated the exclusion of Scytophara extensa from Pereboriidae. Additionally, we estimated divergence times across Pereborioidea lineages, reconstructed ancestral distribution ranges, and elucidated the origin centers and dispersal routes for Curvicubitidae. Wing eyespots and bands suggested predator mimicry, a novel defense strategy in Permian insects.

Keywords

ancestral distribution range / Curvicubitidae / eyespots / new species / phylogenetic relationships / Upper Permian

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Ziqiang Xu, ChungKun Shih, Chenxin Zhou, Wei Yuan, Yu Liu, Dong Ren, Ying Wang. A new Late Permian species of Curvicubitidae illuminates the phylogeny and palaobiogeography of Pereborioidea (Insecta: Hemiptera). Journal of Systematics and Evolution, 2026, 64 (2) : 330-343 DOI:10.1111/jse.70016