Vegetation Patterns: Structures and Dynamics

Li-Feng Hou , Jun Zhang , Gui-Quan Sun , Zhen Jin

CSIAM Trans. Life Sci. ›› 2026, Vol. 2 ›› Issue (1) : 91 -132.

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CSIAM Trans. Life Sci. ›› 2026, Vol. 2 ›› Issue (1) :91 -132. DOI: 10.4208/csiam-ls.SO-2025-0024
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Vegetation Patterns: Structures and Dynamics
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Abstract

Vegetation patterns are a hallmark of ecosystem self-organization, emerg- ing from the intrinsic dynamics of nonlinear feedback mechanisms and spatiotem- poral interactions. This review systematically explores and examines the structural characteristics of these patterns, the phenomena of multistability, and their implica- tions for ecosystem stability through the lens of mathematical modeling and dynam- ical systems theory. In particular, reaction-diffusion models serve as a key analytical tool, revealing how local positive feedback and non-local negative feedback drive self- organized spatial structures via Turing bifurcation. Bifurcation theory and potential landscape analysis further elucidate ecosystem multistability, quantifying critical tran- sitions among uniform vegetation, patterned states, and bare soil under environmen- tal conditions. Advances in spatial metrics, including traditional statistical measures (e.g. variance, autocorrelation) and emerging complexity-based indicators (e.g. hyper- uniformity, spatial permutation entropy) provide robust methods for detecting ecolog- ical functional shifts and early-warning signs of regime shifts. Additionally, restoration strategies grounded in structural optimization, such as optimal control theory, offer a theoretical framework for vegetation pattern reconstruction and stability regulation, particularly in arid and semi-arid regions. Future research should integrate multiscale modeling and interdisciplinary approaches to deepen our understanding of vegetation structure-function relationships. Such efforts will yield both theoretical insights and practical solutions for mitigating global ecological degradation and climate change.

Keywords

Vegetation patterns / ecosystem stability / multistability / optimal control / critical tran-sitions

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Li-Feng Hou, Jun Zhang, Gui-Quan Sun, Zhen Jin. Vegetation Patterns: Structures and Dynamics. CSIAM Trans. Life Sci., 2026, 2(1): 91-132 DOI:10.4208/csiam-ls.SO-2025-0024

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References

[1]

M. Alfaro,H. Izuhara, and M. Mimura, On a nonlocal system for vegetation in drylands, J. Math. Biol., 77:1761-1793, 2018.

[2]

B. Ayhan, C. Kwan, B. Budavari, L. Kwan, Y. Lu, D. Perez, J. Li, D. Skarlatos, and M. Vla-chos, Vegetation detection using deep learning and conventional methods, Remote Sens., 12:2502, 2020.

[3]

R. Bastiaansen, A. Doelman, M. B. Eppinga, and M. Rietkerk, The effect of climate change on the resilience of ecosystems with adaptive spatial pattern formation, Ecol. Lett., 23:414-429, 2020.

[4]

R. Bastiaansen, O. Ja¨ıbi, V. Deblauwe, M. B. Eppinga, K. Siteur, E. Siero, S. Mermoz, A. Bou-vet, A. Doelman, and M. Rietkerk, Multistability of model and real dryland ecosystems through spatial self-organization, Proc. Natl. Acad. Sci. USA, 115:11256-11261, 2018.

[5]

S. Bathiany, D. Nian, M. Dr ¨uke, and N. Boers, Resilience indicators for tropical rainforests in a dynamic vegetation model, Glob. Change Biol., 30:e17613, 2024.

[6]

M. Baudena and M. Rietkerk, Complexity and coexistence in a simple spatial model for arid savanna ecosystems, Theor. Ecol., 6:131-141, 2013.

[7]

J. J. R. Bennett and J. A. Sherratt, Long-distance seed dispersal affects the resilience of banded vegetation patterns in semi-deserts, J. Math. Biol., 481:151-161, 2019.

[8]

M. Berdugo, J. J. Gaita´n, M. Delgado-Baquerizo, T. W. Crowther, and V. Dakos, Preva-lence and drivers of abrupt vegetation shifts in global drylands, Proc. Natl. Acad. Sci. USA, 119:e2123393119, 2022.

[9]

M. Berdugo, S. K´efi,S. Soliveres, and F. T. Maestre, Plant spatial patterns identify alternative ecosystem multifunctionality states in global drylands, Nat. Ecol. Evol., 1:0003, 2017.

[10]

P. N. Bernardino et al., Predictability of abrupt shifts in dryland ecosystem functioning, Nat. Clim. Change, 15:86-91, 2025.

[11]

M. Bianchini et al., Modeling climate-driven vegetation changes under contrasting temperate and arid conditions in the Mediterranean basin, Ecol. Evol., 15:e70753, 2025.

[12]

N. Boers and M. Rypdal,Critical slowing down suggests that the western Greenland ice sheet is close to a tipping point, Proc. Natl. Acad. Sci. USA, 118:e2024192118, 2021.

[13]

F. Borgogno, P. D’odorico, F. Laio, and L. Ridolfi, Mathematical models of vegetation pattern formation in ecohydrology, Rev. Geophys., 47:RG000256, 2009.

[14]

T. M. Bury, R. Sujith, I. Pavithran, M. Scheffer, T. M. Lenton, M. Anand, and C. T. Bauch, Deep learning for early warning signals of tipping points, Proc. Natl. Acad. Sci. USA, 118:e2106140118, 2021.

[15]

J. E. Buxton, J. F. Abrams, C. A. Boulton, N. Barlow, C. Rangel Smith, S. Van Stroud, K. J. Lees, and T. M. Lenton, Quantitatively monitoring the resilience of patterned vegetation in the Sahel, Glob. Change Biol., 28:571-587, 2022.

[16]

S. R. Carpenter and W. A. Brock, Rising variance: A leading indicator of ecological transition, Ecol. Lett., 9:311-318, 2006.

[17]

Z. Chen, P. Fan, X. Hou, F. Ji, L. Li, Z. Qian, G. Feng, and G. Sun, Analysis of global vegetation resilience under different future climate scenarios, Clim. Dyn., 62:7967-7980, 2024.

[18]

Z. Chen, H. Liu, C. Xu, X. Wu, B. Liang, J. Cao, and D. Chen, Modeling vegetation greenness and its climate sensitivity with deep-learning technology, Ecol. Evol., 11:7335-7345, 2021.

[19]

Z. Chen, J. Liu, L. Li, Y. Wu, G. Feng, Z. Qian, and G.-Q. Sun, Effects of climate change on vegetation patterns in Hulun Buir Grassland, Phys. A, 597:127275, 2022.

[20]

Z. Chen, Y.-P. Wu, G.-L. Feng,Z.-H. Qian, and G.-Q. Sun, Effects of global warming on pattern dynamics of vegetation: Wuwei in China as a case, Appl. Math. Comput., 390:125666, 2021.

[21]

C. Ciemer, N. Boers, M. Hirota, J. Kurths, F. M ¨uller-Hansen, R. S. Oliveira, and R. Winkel-mann, Higher resilience to climatic disturbances in tropical vegetation exposed to more variable rainfall, Nat. Geosci., 12:174-179, 2019.

[22]

V. Dakos, S. K´efi, M. Rietkerk, E. H. van Nes, and M. Scheffer, Slowing down in spatially patterned ecosystems at the brink of collapse, Am. Nat., 177:E153-E166, 2011.

[23]

V. Dakos, M. Scheffer, E. H. van Nes, V. Brovkin, V. Petoukhov, and H. Held, Slowing down as an early warning signal for abrupt climate change, Proc. Natl. Acad. Sci. USA, 105:14308-14312, 2008.

[24]

V. Dakos, E. H. van Nes, R. Donangelo, H. Fort, and M. Scheffer, Spatial correlation as leading indicator of catastrophic shifts, Theor. Ecol., 3:163-174, 2010.

[25]

H. de Paoli, T. van der Heide, A. van den Berg, B. R. Silliman, P. M. J. Herman, and J. van de Koppel, Behavioral self-organization underlies the resilience of a coastal ecosystem, Proc. Natl. Acad. Sci. USA, 114:8035-8040, 2017.

[26]

V. Deblauwe, P. Couteron, O. Lejeune, J. Bogaert, and N. Barbier, Environmental modulation of self-organized periodic vegetation patterns in Sudan, Ecography, 34:990-1001, 2011.

[27]

L. Eigentler and J. A. Sherratt, Analysis of a model for banded vegetation patterns in semi-arid environments with nonlocal dispersal, J. Math. Biol., 77:739-763, 2018.

[28]

L. Eigentler and J. A. Sherratt, Long-range seed dispersal enables almost stationary patterns in a model for dryland vegetation, J. Math. Biol., 86:15, 2023.

[29]

M. A. Ferr´e, I. Pavithran, B. K. Bera, H. Uecker, and E. Meron, Vegetation pattern formation and community assembly under drying climate trends, Chaos, 35:093114, 2025.

[30]

U. Feudel, A. N. Pisarchik, and K. Showalter, Multistability and tipping: From mathematics and physics to climate and brain - Minireview and preface to the focus issue, Chaos, 28:033501, 2018.

[31]

B. M. Flores et al., Critical transitions in the Amazon forest system, Nature, 626:555-564, 2024.

[32]

G. Forzieri, V. Dakos, N. G. McDowell, A. Ramdane, and A. Cescatti, Emerging signals of declining forest resilience under climate change, Nature, 608:534-539, 2022.

[33]

Z. Ge,The hidden order of Turing patterns in arid and semi-arid vegetation ecosystems, Proc. Natl. Acad. Sci. USA, 120:e2306514120, 2023.

[34]

S. Getzin et al., Discovery of fairy circles in Australia supports self-organization theory, Proc. Natl. Acad. Sci. USA, 113:3551-3556, 2016.

[35]

E. Gilad, J. von Hardenberg, A. Provenzale, M. Shachak, and E. Meron, Ecosystem engineers: From pattern formation to habitat creation, Phys. Rev. Lett., 93:098105, 2004.

[36]

E. Gilad, J. von Hardenberg, A. Provenzale,M. Shachak, and E. Meron, A mathematical model of plants as ecosystem engineers, J. Theor. Biol., 244:680-691, 2007.

[37]

P. Gonzalez, R. P. Neilson,J. M. Lenihan, and R. J. Drapek, Global patterns in the vulnerability of ecosystems to vegetation shifts due to climate change, Glob. Ecol. Biogeogr., 19:755-768, 2010.

[38]

V. Guttal and C. Jayaprakash, Changing skewness: An early warning signal of regime shifts in ecosystems, Ecol. Lett., 11:450-460, 2008.

[39]

V. Guttal and C. Jayaprakash, Spatial variance and spatial skewness: Leading indicators of regime shifts in spatial ecological systems, Theor. Ecol., 2:3-12, 2009.

[40]

S. I. Higgins and S. Scheiter, Atmospheric CO 2 forces abrupt vegetation shifts locally, but not globally, Nature, 488:209-212, 2012.

[41]

R. HilleRisLambers, M. Rietkerk, F. van den Bosch, H. H. T. Prins, and H. de Kroon, Vege-tation pattern formation in semi-arid grazing systems, Ecology, 82:50-61, 2001.

[42]

M. Hirota, M. Holmgren, E. H. van Nes, and M. Scheffer, Global resilience of tropical forest and savanna to critical transitions, Science, 334:232-235, 2011.

[43]

L.-F. Hou, S.-P. Gao, L.-L. Chang, Y.-P. Wu, G.-L. Feng, Z. Wang, and G.-Q. Sun, Vegetation restoration strategies in arid or semi-arid regions - From the perspective of optimal control, Chaos, 34:113109, 2024.

[44]

L.-F. Hou, S.-P. Gao, and G.-Q. Sun, Two types of fairy circles coexist in a vegetation - water model, Nonlinear Dyn., 111:7883-7898, 2023.

[45]

L.-F. Hou, L. Li, L. Chang, Z. Wang, and G.-Q. Sun, Pattern dynamics of vegetation based on optimal control theory, Nonlinear Dyn., 113:1-23, 2025.

[46]

L.-F. Hou, L. Li, R. Chen, Y.-P. Wu, G.-L. Feng, and G.-Q. Sun, Vegetation dynamics: Modeling, mechanisms, and emergent properties, Phys. Rep., 1145:1-87, 2025.

[47]

L.-F. Hou,G.-Q. Sun, and M. Perc, The impact of heterogeneous human activity on vegetation patterns in arid environments, Commun. Nonlinear Sci. Numer. Simul., 126:107461, 2023.

[48]

W. Hu, L. Cui, M. Delgado-Baquerizo, R. Sol´e, S. K´efi, M. Berdugo, N. Xu, B. Wang, Q.-X. Liu, and C. Xu, Causes and consequences of disordered hyperuniformity in global drylands, Proc. Natl. Acad. Sci. USA, 122:e2504496122, 2025.

[49]

Inderjit R. M. Callaway and E. Meron, Belowground feedbacks as drivers of spatial self-organization and community assembly, Phys. Life Rev., 38:1-24, 2021.

[50]

L. Jiang, G. Jiapaer, A. Bao, H. Guo, and F. Ndayisaba, Vegetation dynamics and responses to climate change and human activities in Central Asia, Sci. Total Environ., 599:967-980, 2017.

[51]

S. K´efi, A. G´enin, A. Garcia-Mayor, E. Guirado, J. S. Cabral, M. Berdugo, J. Guerber, R. Sol´e, and F. T. Maestre, Self-organization as a mechanism of resilience in dryland ecosystems, Proc. Natl. Acad. Sci. USA, 121:e2305153121, 2024.

[52]

S. K´efi, V. Guttal, W. A. Brock, S. R. Carpenter, A. M. Ellison, V. N. Livina, D. A. Seekell, M. Scheffer, E. H. van Nes, and V. Dakos, Early warning signals of ecological transitions: Meth-ods for spatial patterns, PLoS ONE, 9:e92097, 2014.

[53]

S. K´efi, M. Rietkerk, C. Alados, Y. Pueyo, V. P. Papanastasis, A. ElAich, and P. C. de Ruiter, Spatial vegetation patterns and imminent desertification in Mediterranean arid ecosystems, Na-ture, 449:213-217, 2007.

[54]

S. Kefi,M. Rietkerk, and G. G. Katul, Vegetation pattern shift as a result of rising atmospheric CO2 in arid ecosystems, Theor. Popul. Biol., 74:332-344, 2008.

[55]

C. A. Klausmeier, Regular and irregular patterns in semiarid vegetation, Science, 284:1826-1828, 1999.

[56]

C. Kuehn, A mathematical framework for critical transitions: Normal forms, variance and applica-tions, J. Nonlinear Sci., 23:457-510, 2013.

[57]

S. Lavorel, Ecological diversity and resilience of mediterranean vegetation to disturbance, Divers. Distrib., 5:3-13, 1999.

[58]

R. Lefever and O. Lejeune, On the origin of tiger bush, Bull. Math. Biol., 59:263-294, 1997.

[59]

J. Li,G.-Q. Sun, and Z. Jin, Interactions of time delay and spatial diffusion induce the periodic oscillation of the vegetation system, Discrete Contin. Dyn. Syst. Ser. B, 27:2147-2172, 2022.

[60]

J. Li, G.-Q. Sun, L. Li,Z. Jin, and Y. Yuan, The effect of grazing intensity on pattern dynamics of the vegetation system, Chaos Solit. Fractals, 175:114025, 2023.

[61]

L. Li, Y.-Z. Pang, G.-Q. Sun, and S. Ruan, Impact of climate change on vegetation patterns in Altay Prefecture, China, Math. Med. Biol., 41:53-80, 2024.

[62]

J. Liang and G.-Q. Sun, Effects of climate change on vegetation pattern in Baotou, China, Non-linear Dyn., 112:8675-8693, 2024.

[63]

Y. Lin, G. Han, M. Zhao, and S. X. Chang, Spatial vegetation patterns as early signs of deserti-fication: A case study of a desert steppe in Inner Mongolia, China, Landsc. Ecol., 25:1519-1527, 2010.

[64]

Z. Liu, X. Zhang, X. Ru, T.-T. Gao, J. M. Moore, and G. Yan, Early predictor for the onset of critical transitions in networked dynamical systems, Phys. Rev. X., 14:031009, 2024.

[65]

M. T. Lo¨bmann, C. Geitner, C. Wellstein, and S. Zerbe, The influence of herbaceous vegetation on slope stability - A review, Earth-Sci. Rev., 209:103328, 2020.

[66]

M. Lo´pez Pereira, V. O. Sadras, W. Batista, J. J. Casal, and A. J. Hall, Light-mediated self-organization of sunflower stands increases oil yield in the field, Proc. Natl. Acad. Sci. USA, 114:7975-7980, 2017.

[67]

S. Ma, J. Ren, C. Wu, and Q. He, Extreme precipitation events trigger abrupt vegetation succes-sion in emerging coastal wetlands, Catena, 241:108066, 2024.

[68]

F. T. Maestre and A. Escudero, Is the patch size distribution of vegetation a suitable indicator of desertification processes?, Ecology, 90:1729-1735, 2009.

[69]

R. Mart´ınez-Garc´ıa, J. M. Calabrese, E. Herna´ndez-Garc´ıa, and C. Lo´pez, Vegetation pattern formation in semiarid systems without facilitative mechanisms, Geophys. Res. Lett., 40:6143-6147, 2013.

[70]

R. Mart´ınez-Garc´ıa,J. M. Calabrese, and C. Lo´pez, Spatial patterns in mesic savannas: The local facilitation limit and the role of demographic stochasticity, J. Theor. Biol., 333:156-165, 2013.

[71]

Y. Mau, A. Hagberg, and E. Meron, Spatial periodic forcing can displace patterns it is intended to control, Phys. Rev. Lett., 109:034102, 2012.

[72]

Y. Mau, L. Haim and E. Meron, Reversing desertification as a spatial resonance problem, Phys. Rev. E, 91:012903, 2015.

[73]

E. Meron, Pattern-formation approach to modelling spatially extended ecosystems, Ecol. Model., 234:70-82, 2012.

[74]

E. Meron, Pattern formation - A missing link in the study of ecosystem response to environmental changes, Math Biosci., 271:1-18, 2016.

[75]

E. Meron, From patterns to function in living systems: Dryland ecosystems as a case study, Annu. Rev. Condens. Matter Phys., 9:79-103, 2018.

[76]

E. Meron, Vegetation pattern formation: The mechanisms behind the forms, Phys. Today, 72:30-36, 2019.

[77]

E. Meron, E. Gilad, J. Von Hardenberg,M. Shachak, and Y. Zarmi, Vegetation patterns along a rainfall gradient, Chaos Solit. Fractals, 19:367-376, 2004.

[78]

W. Pałubicki, M. Makowski, W. Gajda, T. Ha¨drich,D. L. Michels, and S. Pirk, Ecoclimates: Climate-response modeling of vegetation, ACM Trans. Graph., 41:1-19, 2022.

[79]

Y.-Z. Pang, L. Li, and Z. Jin, Early warning signals of critical transitions in ecosystems: Entropy reduction in vegetation spatial patterns, Nonlinear Dyn., 113:15597-15618, 2025.

[80]

B. Pichon, I. Gounand, S. Donnet, and S. K´efi, The interplay of facilitation and competition drives the emergence of multistability in dryland plant communities, Ecology, 105:e4369, 2024.

[81]

Y. Pueyo, S. K´efi, C. Alados, and M. Rietkerk, Dispersal strategies and spatial organization of vegetation in arid ecosystems, Oikos, 117:1522-1532, 2008.

[82]

M. Reichstein, M. Bahn, M. D. Mahecha, J. Kattge, and D. D. Baldocchi, Linking plant and ecosystem functional biogeography, Proc. Natl. Acad. Sci. USA, 111:13697-13702, 2014.

[83]

M. Rietkerk, R. Bastiaansen, S. Banerjee, J. van de Koppel, M. Baudena, and A. Doelman, Evasion of tipping in complex systems through spatial pattern formation, Science, 374:eabj0359, 2021.

[84]

M. Rietkerk, M. C. Boerlijst, F. van Langevelde, R. HilleRisLambers, J. van de Koppel, L. Kumar, H. H. T. Prins, and A. M. de Roos, Self-organization of vegetation in arid ecosys-tems, Am. Nat., 160:524-530, 2002.

[85]

M. Rietkerk, S. C. Dekker, P. C. de Ruiter, and J. van de Koppel, Self-organized patchiness and catastrophic shifts in ecosystems, Science, 305:1926-1929, 2004.

[86]

M. Rietkerk and J. van de Koppel, Regular pattern formation in real ecosystems, Trends Ecol. Evol., 23:169-175, 2008.

[87]

M. Scheffer, Critical Transitions in Nature and Society, in: Princeton Studies in Complexity, Vol. 16, Princeton University Press, 2020.

[88]

M. Scheffer, J. Bascompte, W. A. Brock, V. Brovkin, S. R. Carpenter, V. Dakos, H. Held, E. H. van Nes, M. Rietkerk, and G. Sugihara, Early-warning signals for critical transitions, Nature, 461:53-59, 2009.

[89]

M. Scheffer and S. R. Carpenter, Catastrophic regime shifts in ecosystems: Linking theory to observation, Trends Ecol. Evol., 18:648-656, 2003.

[90]

M. Scheffer, S. Carpenter, J. A. Foley, C. Folke, and B. Walker, Catastrophic shifts in ecosystems, Nature, 413:591-596, 2001.

[91]

M. Scheffer, S. R. Carpenter, T. M. Lenton, J. Bascompte, W. Brock, V. Dakos, J. van de Koppel, I. A. van de Leemput, S. A. Levin, E. H. van Nes, M. Pascual, and J. Vandermeer, Anticipating critical transitions, Science, 338:344-348, 2012.

[92]

F. Schweisguth and F. Corson, Self-organization in pattern formation Dev. Cell, 49:659-677, 2019.

[93]

J. A. Sherratt, An analysis of vegetation stripe formation in semi-arid landscapes, J. Math. Biol., 51:183-197, 2005.

[94]

J. A. Sherratt, Pattern solutions of the Klausmeier model for banded vegetation in semi-arid envi-ronments I, Nonlinearity, 23:2657, 2010.

[95]

J. A. Sherratt, Pattern solutions of the Klausmeier model for banded vegetation in semiarid en-vironments IV: Slowly moving patterns and their stability, SIAM J. Appl. Math., 73:330-350, 2013.

[96]

J. A. Sherratt, Pattern solutions of the Klausmeier model for banded vegetation in semiarid envi-ronments V: The transition from patterns to desert, SIAM J. Appl. Math., 73:1347-1367, 2013.

[97]

J. A. Sherratt,Using wavelength and slope to infer the historical origin of semiarid vegetation bands, Proc. Natl. Acad. Sci. USA, 112:4202-4207, 2015.

[98]

E. Siero, A. Doelman, M. B. Eppinga, J. D. M. Rademacher, M. Rietkerk, and K. Siteur, Striped pattern selection by advective reaction-diffusion systems: Resilience of banded vegetation on slopes, Chaos, 25:036411, 2015.

[99]

T. Smith and N. Boers, Global vegetation resilience linked to water availability and variability, Nat. Commun., 14:498, 2023.

[100]

T. Smith,D. Traxl, and N. Boers, Empirical evidence for recent global shifts in vegetation re-silience, Nat. Clim. Change, 12:477-484, 2022.

[101]

G.-Q. Sun, R. He, L.-F. Hou, X. Luo, S. Gao, L. Chang, Y. Wang, and Z.-K. Zhang, Optimal control of spatial diseases spreading in networked reaction-diffusion systems, Phys. Rep., 1111:1-64, 2025.

[102]

G.-Q. Sun, L.-F. Hou, L. Li,Z. Jin, and H. Wang, Spatial dynamics of a vegetation model with uptake - diffusion feedback in an arid environment, J. Math. Biol., 85:50, 2022.

[103]

G.-Q. Sun, L. Li, J. Li, C. Liu, Y.-P. Wu, S. Gao, Z. Wang, and G.-L. Feng, Impacts of climate change on vegetation pattern: Mathematical modeling and data analysis, Phys. Life Rev., 43:239-270, 2022.

[104]

G.-Q. Sun, C.-H. Wang, L.-L. Chang, Y.-P. Wu,L. Li, and Z. Jin, Effects of feedback regulation on vegetation patterns in semi-arid environments, Appl. Math. Model., 61:200-215, 2018.

[105]

G.-Q. Sun, Z.-C. Xue, L. Li, J. Li, C. I. del Genio, and S. Boccaletti, Chimera states with multiple coexisting solutions, Phys. Rev. Res., 7:023289, 2025.

[106]

G.-Q. Sun, H.-T. Zhang, Y.-L. Song, L. Li, and Z. Jin, Dynamic analysis of a plant-water model with spatial diffusion, J. Differential Equations, 329:395-430, 2022.

[107]

G.-Q. Sun, H.-T. Zhang, J.-S. Wang, J. Li, Y. Wang, L. Li, Y.-P. Wu, G.-L. Feng, and Z. Jin, Mathematical modeling and mechanisms of pattern formation in ecological systems: A review, Non-linear Dyn., 104:1677-1696, 2021.

[108]

M. P. Thakur, W. H. van der Putten, R. A. Wilschut, G. C. Veen, P. Kardol, J. van Ruijven, E. Allan, C. Roscher,M. van Kleunen, and T. M. Bezemer, Plant-soil feedbacks and temporal dynamics of plant diversity-productivity relationships, Trends Ecol. Evol., 36:651-661, 2021.

[109]

C. Tian,Z. Ling, and L. Zhang, Delay-driven spatial patterns in a network-organized semiarid vegetation model, Appl. Math. Comput., 367:124778, 2020.

[110]

G. Tirabassi and C. Masoller,Entropy-based early detection of critical transitions in spatial veg-etation fields, Proc. Natl. Acad. Sci. USA, 120:e2215667120, 2023.

[111]

S. Torquato, Hyperuniform states of matter, Phys. Rep., 745:1-95, 2018.

[112]

S. Torquato and F. H. Stillinger, Local density fluctuations, hyperuniformity and order metrics, Phys. Rev. E, 68:041113, 2003.

[113]

F. Tro¨ltzsch, Optimal control of partial differential equations: Theory, methods, and applications, in: Graduate Studies in Mathematics, Vol. 112, AMS, 2010.

[114]

C. Valentin, J.-M. d’Herb`es and J. Poesen, Soil and water components of banded vegetation patterns, Catena, 37:1-24, 1999.

[115]

J. van Belzen, J. van de Koppel, M. L. Kirwan, D. van der Wal, P. M. Herman, V. Dakos, S. K´efi, M. Scheffer, G. R. Guntenspergen, and T. J. Bouma, Vegetation recovery in tidal marshes reveals critical slowing down under increased inundation, Nat. Commun., 8:15811, 2017.

[116]

W. H. van der Putten et al., Plant-soil feedbacks: The past, the present and future challenges, Journal of Ecology, 101:265-276, 2013.

[117]

E. H. van Nes and M. Scheffer, Slow recovery from perturbations as a generic indicator of a nearby catastrophic shift, Am. Nat., 169:738-747, 2007.

[118]

J. von Hardenberg, E. Meron, M. Shachak, and Y. Zarmi, Diversity of vegetation patterns and desertification, Phys. Rev. Lett., 87:198101, 2001.

[119]

C. Wang,H. Wang, and S. Yuan, Precipitation governing vegetation patterns in an arid or semi-arid environment, J. Math. Biol., 87:22, 2023.

[120]

X. Wang,J. Shi, and G. Zhang, Bifurcation and pattern formation in an activator-inhibitor model with non-local dispersal, Bull. Math. Biol., 84:140, 2022.

[121]

E. Weerman, J. Van Belzen, M. Rietkerk, S. Temmerman, S. K´efi, P. Herman, and J. Van de Koppel, Changes in diatom patch-size distribution and degradation in a spatially self-organized intertidal mudflat ecosystem, Ecology, 93:608-618, 2012.

[122]

D. Wu, X. Zhao, S. Liang, T. Zhou, K. Huang, B. Tang, and W. Zhao, Time-lag effects of global vegetation responses to climate change, Glob. Change Biol., 21:3520-3531, 2015.

[123]

Q. Xue, C. Liu, L. Li,G.-Q. Sun, and Z. Wang, Interactions of diffusion and nonlocal delay give rise to vegetation patterns in semi-arid environments, Appl. Math. Comput., 399:126038, 2021.

[124]

Z.-C. Xue, J. Li, C.-H. Wang,G.-Q. Sun, and L. Li, Multistability shifts in an aird vegetation system with nonlocal water absorption effect, J. Math. Biol., 91:17, 2025.

[125]

L. Yang, Q. Guan, J. Lin, J. Tian,Z. Tan, and H. Li, Evolution of NDVI secular trends and responses to climate change: A perspective from nonlinearity and nonstationarity characteristics, Remote Sens. Environ., 254:112247, 2021.

[126]

Y. Yang, K. R. Foster,K. Z. Coyte, and A. Li, Time delays modulate the stability of complex ecosystems, Nat. Ecol. Evol., 7:1610-1619, 2023.

[127]

Y. R. Zelnik, P. Gandhi, E. Knobloch, and E. Meron, Implications of tristability in pattern-forming ecosystems, Chaos, 28:033609, 2018.

[128]

Y. R. Zelnik, S. Kinast, H. Yizhaq, G. Bel and E. Meron, Regime shifts in models of dryland vegetation, Philos. Trans. Roy. Soc. A, 371:20120358, 2013.

[129]

Y. R. Zelnik, Y. Mau,M. Shachak, and E. Meron, High-integrity human intervention in ecosys-tems: Tracking self-organization modes, PLoS Comput. Biol., 17:e1009427, 2021.

[130]

Y. R. Zelnik, E. Meron and G. Bel, Gradual regime shifts in fairy circles, Proc. Natl. Acad. Sci. USA, 112:12327-12331, 2015.

[131]

H.-T. Zhang, Y.-P. Wu, G.-Q. Sun,C. Liu, and G.-L. Feng, Bifurcation analysis of a spatial vegetation model, Appl. Math. Comput., 434:127459, 2022.

[132]

L.-X. Zhao, C. Xu, Z.-M. Ge, J. Van De Koppel, and Q.-X. Liu, The shaping role of self-organization:Linking vegetation patterning, plant traits and ecosystem functioning, Proc. R. Soc. B: Biol. Sci., 286:20182859, 2019.

[133]

Q. Zhu et al., An early warning signal for grassland degradation on the Qinghai-Tibetan Plateau, Nat. Commun., 14:6406, 2023.

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