Guangdong Key Laboratory for Genome Stability & Disease Prevention and Carson International Cancer Center, Marshall Laboratory of Biomedical Engineering, Shenzhen University Medical School, Shenzhen University, Shenzhen, Guangdong, China.

Shenzhen Key Laboratory for Systemic Aging and Intervention (SKL-SAI), School of Basic Medical Sciences, Shenzhen University Medical School, Shenzhen, Guangdong, China.

Department of Immunology, University of Tübingen, Tübingen, Germany.

Departments of Biology and Medicine, University of Rochester, Rochester, NY, USA.

Department of Human Genetics, David Geffen School of Medicine, University of California, Los Angeles, CA, USA.
Altos Labs, San Diego, CA, USA. Altos Labs, Cambridge, UK.

Department of Microbiology and Molecular Genetics, University of California, Davis, CA, USA.

University of Chinese Academy of Sciences, Beijing, China.

Department of Epigenetics, Van Andel Institute, Grand Rapids, MI, USA.

State Key Laboratory of Radiation Medicine and Protection and Institutes for Translational Medicine at the First and Third Affiliated Hospitals of Soochow University, Suzhou Medical College of Soochow University, Suzhou, Jiangsu, China.

Department of Medicine, Center for Advanced Gerotherapeutics, Cedars-Sinai Medical Center, Los Angeles, CA, USA.

State Key Laboratory of Microbial Technology, Shandong University, Qingdao, Shandong, China.
Faculty of Biological Sciences, Friedrich-Schiller-University Jena, Jena, Germany
Leibniz Institute on Aging-Fritz Lipmann Institute (FLI), Jena, Germany.

School of Biomedical Sciences, University of Hong Kong, Hong Kong, China.

Institute for Research on Cancer and Aging of Nice (IRCAN), INSERM, Université Côte d'Azur, Centre National de la Recherche Scientifique(CNRS), Nice, France.

Life Sciences Institute, Zhejiang University, Hangzhou, Zhejiang, China.

Department of Oncology, Mayo Clinic, Rochester, MA, USA.

Tongji University School of Medicine, Shanghai, China.

Department of Medicine, Breast Unit, The Royal Marsden NHS Foundation Trust, Sutton, UK.

Cellular senescence, Cancer & Aging Lab, Cancer Research Center of Lyon, Lyon, France.

Institut Curie, PSL Research University, CNRS UMR3244, Dynamics of Genetic Information, Sorbonne Université, Paris, France.

Shanghai Key Laboratory for Tumor Microenvironment and Inflammation, Department of Biochemistry and Molecular Cell Biology, Shanghai Jiao Tong University School of Medicine, Shanghai, China. Institute of Aging & Tissue Regeneration, Ren-Ji Hospital, Shanghai Jiao Tong University School of Medicine, Shanghai, China. Hainan Academy of Medical Sciences, Haikou, Hainan, China.

Gustave Roussy, Université Paris-Saclay, Villejuif, France.

IFOM, The AIRC Institute of Molecular Oncology, Milan, Italy.

National Clinical Research Center for Geriatrics, State Key Laboratory of Biotherapy, West China Hospital, Sichuan University, Chengdu, Sichuan, China.

Department of Oncology, MZB Foundation for Cancer Research, New York, NY, USA.

Geriatric Unit, Lyon Sud Hospital, Pierre-Benite, France.

Rutgers Cancer Institute, Rutgers University, New Brunswick, NJ, USA.

Université Paris Cité, INSERM U970, PARCC, Paris, France.

Key Laboratory of Regenerative Medicine of Ministry of Education, Institute of Aging and Regenerative Medicine, College of Life Science and Technology, Jinan University, Guangzhou, Guangdong, China.

Department of Molecular Biology, Princeton University, Princeton, NJ, USA.

Cordeliers Research Center, Université de Paris Cité, Paris, France.

Department of Medicine, Faculty of Medicine and Health Sciences, University of Sherbrooke, Sherbrooke, QC, Canada.

Department of Dermatology, Duke University School of Medicine, Durham, NC, USA.
Institute for Molecular and Cellular Therapeutics, Chinese Institutes for Medical Research, Beijing, China. School of Basic Medical Sciences, Capital Medical University, Beijing, China.

The First Affiliated Hospital, School of Medicine, Zhejiang University, Hangzhou, Zhejiang, China.

Department of Urology, Johns Hopkins University, Baltimore, MD, USA.

Department of Immunology, Leiden University Medical Center, Oegstgeest, The Netherlands.

Center for Genomic Integrity, Institute for Basic Science, Ulsan National Institute of Science and Technology, Ulsan, South Korea.

Department of Hematology and Stem Cell Transplantation, University Hospital Essen, University Duisburg-Essen, Hufelandstrasse, Essen, Germany.

Department of Medical Oncology, Leiden University Medical Center, Leiden, The Netherlands.

Key Laboratory of Organ Regeneration and Reconstruction, State Key Laboratory of Membrane Biology, State Key Laboratory of Stem Cell and Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China.

State Key Laboratory of Radiation Medicine and Protection, Institutes for Translational Medicine, Soochow University, Suzhou, Jiangsu, China.

Department of Immunology, Universidad De Cordoba, Cordoba, Spain.

Department of Physiology, Immunology Unit, University of Extremadura, Cáceres, Spain.

Department of Biochemistry, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.

Department of hematology, oncology and immunotherapy cantonal hospital Baselland, Medical University Clinics, Basel, Switzerland.

Department of Biomedical Engineering, Faculty of Engineering, Hong Kong Polytechnic University, Hong Kong, China.

School of Life Sciences, Sichuan University, Chengdu, Sichuan, China.

State Key Laboratory of Chemical Oncogenomics, Shenzhen International Graduate School, Tsinghua University, Shenzhen, Guangdong, China. Shenzhen Key Laboratory of Health Science and Technology, Institute of Biopharmaceutical and Health, Shenzhen International Graduate School, Tsinghua University, Shenzhen, Guangdong, China. Open FIESTA Center, Shenzhen International Graduate School, Tsinghua University, Shenzhen, Guangdong, China.

Westlake Laboratory of Life Sciences and Biomedicine, Hangzhou, Zhejiang, China.

The Key Laboratory for Stem Cells and Tissue Engineering, The First Affiliated Hospital, and Department of Histology and Embryology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, China.

Suzhou Institute of Systems Medicine, Chinese Academy of Medical Sciences & Peking Union Medical College, Suzhou, Jiangsu, China.

China National Center for Bioinformation, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China.